1. ARC Centre of Excellence for Plant Success in Nature and Agriculture, The University of Queensland, St Lucia, QLD 4072, Australia
2. School of Biological Sciences, The University of Queensland, St Lucia, QLD 4072, Australia
3. Centre for Horticultural Science, Queensland Alliance for Agriculture and Food Innovation, The University of Queensland, Brisbane, QLD 4068, Australia
4. J&S Plant Biotech, P.O. Box 4700, Sunshine Coast MC, QLD 4560, Australia