1. College of Veterinary Medicine, Qingdao Agricultural University, Qingdao 266109, China
2. College of Life Science, Qingdao Agricultural University, Qingdao 266109, China
3. The Key Laboratory of Experimental Marine Biology, Centre for Ocean Mega-Science, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
4. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266071, China
5. Jinan Center for Animal Disease Prevention and Control, Jinan 250109, China
6. Southern Marine Science and Engineering Guangdong Laboratory, Guangzhou 511458, China