1. Industry-University Cooperation Foundation, Pukyong National University, 45 Yongso-ro, Nam-Gu, Busan 48513, Republic of Korea
2. Department of Microbiology, Pukyong National University, 45 Yongso-ro, Nam-Gu, Busan 48513, Republic of Korea
3. School of Marine and Fisheries Life Science, Pukyong National University, 45 Yongso-ro, Nam-Gu, Busan 48513, Republic of Korea
4. Department of Marine Biology, Pukyong National University, 45 Yongso-ro, Nam-Gu, Busan 48513, Republic of Korea
5. Research Institute for Basic Science, Pukyong National University, 45 Yongso-ro, Nam-Gu, Busan 48513, Republic of Korea
6. Gladstone Institute, San Francisco, CA 94158, USA