1. Ecosystem Science and Management, Penn State University, 432 Forest Resources Building, University Park, PA 16802, USA
2. South African Institute for Aquatic Biodiversity, Makhanda 6140, South Africa
3. School of Forest Resources, Penn State University, University Park, PA 16802, USA
4. Pennsylvania Fish and Boat Commission, 595 Rolling Ridge Drive, Bellefonte, PA 16823, USA
5. Pennsylvania Game Commission, Harrisburg, PA 19019, USA