1. Division of Forestry and Natural Resources, West Virginia University, 333 Evansdale Drive, Morgantown, WV 26506, USA
2. U.S. Geological Survey, Eastern Ecological Science Center at Leetown Research Laboratory, 11649 Leetown Road, Kearneysville, WV 25430, USA
3. U.S. Geological Survey, West Virginia Cooperative Fish and Wildlife Research Unit, West Virginia University, Morgantown, WV 26506, USA