1. Environmental & Conservation Sciences Graduate Program, North Dakota State University, Fargo, ND 58102, USA
2. Department of Biosciences, Minnesota State University Moorhead, Moorhead, MN 56563, USA
3. Arrowhead Ecology Group, 110 Bugg Lane, San Marcos, TX 78667, USA
4. Department of Biological Sciences, North Dakota State University, Fargo, ND 58102, USA