1. Department of Biology, College of Sciences, San Diego State University, San Diego, CA 92182, USA
2. Center for Complex Human-Environment Relationships, San Diego State University, San Diego, CA 92182, USA
3. Department of Landscape Architecture, College of Design Construction and Planning, University of Florida, Gainesville, FL 32601, USA
4. Department of Geography, College of Arts and Letters, San Diego State University, San Diego, CA 92182, USA
5. College of Forestry, Wildlife and Environment, Auburn University, Auburn, AL 36849, USA