1. Pyrenean Institute of Ecology (CSIC), 22700 Jaca, Spain
2. Institute for Game and Wildlife Research IREC (CSIC-UCLM-JCCM), 13071 Ciudad Real, Spain
3. Cos d’Agents Rurals, Àrea Bàsica del Pallars Jussà, 25620 Tremp, Spain
4. Department of Evolutionary Ecology, Estación Biológica de Doñana (CSIC), 41092 Sevilla, Spain