1. Department of Immunology, Tufts University School of Medicine, 136 Harrison Avenue, Boston, MA 02111, USA
2. Program in Cell, Molecular and Developmental Biology, Graduate School of Biomedical Sciences, Tufts University, 136 Harrison Avenue, Boston, MA 02111, USA
3. Program in Pharmacology, Graduate School of Biomedical Sciences, Tufts University, 136 Harrison Avenue, Boston, MA 02111, USA
4. Program in Immunology, Graduate School of Biomedical Sciences, Tufts University, 136 Harrison Avenue, Boston, MA 02111, USA
5. Department of Orthopaedics, Tufts Medical Center, 800 Washington Street, Boston, MA 02111, USA