1. Department of Biodiversity and Evolutionary Biology, Museo Nacional de Ciencias Naturales (MNCN-CSIC), C/José Gutiérrez Abascal, 2, 28006 Madrid, Spain
2. Department of Biodiversity, Ecology and Evolution, Universidad Complutense de Madrid, C/José Antonio Novais, 12, 28040 Madrid, Spain
3. Real Jardín Botánico (RJB-CSIC), Pl. Murillo, 2, 28014 Madrid, Spain