1. Research College of Forestry, Beijing Forestry University, Beijing 100083, China
2. Faculty of Forestry and Forest Ecology, Georg-August-University, 37077 Göttingen, Germany
3. Department of Forest and Wood Science, Stellenbosch University, Stellenbosch 7602, South Africa
4. State Key Laboratory of Efficient Production of Forest Resources, College of Forestry, Beijing Forestry University, Beijing 100083, China