1. Animal Production, Agricultural Research Council, Private Bag X 02, Irene 0062, South Africa
2. School of Life Sciences, University of KwaZulu-Natal, Scottsville 3209, South Africa
3. Animal and Poultry Science, School of Agriculture, Earth and Environmental Sciences, University of KwaZulu-Natal, Scottsville 3209, South Africa
4. Department of Biological Sciences, Kent State University, Cunningham Hall, Kent, OH 44242, USA