1. School of Life Sciences, Guizhou Normal University, Guiyang 550025, China
2. Eastern China Conservation Centre for Wild Endangered Plant Resources, Shanghai Chenshan Botanical Garden, Shanghai 201602, China
3. College of Forestry, Hainan University, Haikou 570228, China
4. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100000, China
5. China National Botanical Garden, Beijing 100000, China