1. Department of Plant Science, University of Manitoba, Winnipeg, MB R3T 2N2, Canada
2. Department of Botany, Faculty of Science, Tanta University, Tanta 31527, Egypt
3. Department of Genetics, Faculty of Agriculture, University of Tanta, Tanta 31527, Egypt
4. Agriculture and Agri-Food Canada, Ottawa, ON K1A 0C6, Canada