1. Research Department, Siberian Federal University, 660041 Krasnoyarsk, Russia
2. Institute of Fundamental Biology and Biotechnology, Siberian Federal University, 660041 Krasnoyarsk, Russia
3. Institute of Ecology and Geography, Siberian Federal University, 660041 Krasnoyarsk, Russia
4. Khakass Technical Institute, Siberian Federal University, 655017 Abakan, Russia
5. College of Life Sciences, Zhejiang University, Hangzhou 310058, China
6. School of Geography and Ocean Science, Nanjing University, Nanjing 210023, China