1. Key Laboratory of Marine Genetics and Breeding (OUC), Ministry of Education, Qingdao 266000, China
2. College of Marine Life Sciences, Ocean University of China, Qingdao 266000, China
3. Key Laboratory of Utilization and Conservation for Tropical Marine Bioresources (Hainan Tropical Ocean University), Ministry of Education, Sanya 572000, China
4. Department of Plant Biochemistry, Ruhr University Bochum, 44787 Bochum, Germany