1. Sanya Nanfan Research Institute of Hainan University, Hainan Yazhou Bay Seed Laboratory, Sanya 572025, China
2. College of Tropical Crops, Hainan University, Haikou 570228, China
3. Rubber Research Institute, Chinese Academy of Tropical Agricultural Science, Haikou 571101, China
4. Haikou Key Laboratory of Innovation of Seedlings of Tropical Plants, Haikou 571101, China