1. Zhenjiang Academy of Agricultural Sciences, Zhenjiang Institute of Agricultural Sciences in Hilly Areas of Jiangsu Province, Zhenjiang 212400, China
2. The Engineering Research Institute of Agriculture and Forestry, Ludong University, No. 186 Hongqizhong Road, Yantai 264025, China
3. Faculty of Modern Agriculture, Linyi Vocational University of Science and Technology, No. 1 Macau Road, Linyi 276000, China
4. Department of Plant Science, University of Cambridge, Cambridge CB2 3EA, UK
5. Wolfson College, University of Cambridge, Cambridge CB3 9BB, UK