1. Center for Ecological Research, Northeast Forestry University, Harbin 150040, China
2. Key Laboratory of Sustainable Forest Ecosystem Management-Ministry of Education, Northeast Forestry University, Harbin 150040, China
3. Northeast Asia Biodiversity Research Center, Northeast Forestry University, Harbin 150040, China
4. CAS Key Laboratory of Forest Ecology and Management, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110164, China
5. Heilongjiang Forestry Engineering and Environment Institute, Harbin 150040, China
6. Key Laboratory of Forest Ecology and Forestry Ecological Engineering of Heilongjiang Province, Harbin 150040, China