1. Department of Pharmacy, Guizhou University of Traditional Chinese Medicine, Guiyang 550025, China
2. Plant Germplasm and Genomics Center, Germplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China
3. Department of Biology, College of Letters and Sciences, Columbus State University, University System of Georgia, Columbus, GA 31907–5645, USA
4. Lijiang Forest Biodiversity National Observation and Research Station, Kunming Institute of Botany, Chinese Academy of Sciences, Lijiang 674100, China