1. College of Landscape Architecture and Life Science, Chongqing University of Arts and Sciences, Chongqing 402160, China
2. College of Horticulture and Gardening, Yangtze University, Jingzhou 433200, China
3. College of Biology and Food Engineering, Chongqing Three Gorges University, Chongqing 404100, China
4. Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China