1. College of Life Sciences, Shanxi Agricultural University, Taigu 030810, China
2. Shanxi Key Laboratory of Grassland Ecological Protection and Native Grass Germplasm Innovation, College of Grassland Science, Shanxi Agricultural University, Taigu 030801, China
3. Youyu Loess Plateau Grassland Ecosystem Research Station, Shanxi Agricultural University, Taigu 030801, China
4. State Key Laboratory of Vegetation and Environmental Change, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China