1. Grassland College, Xinjiang Agricultural University, Urumqi 830011, China
2. CAS Key Laboratory of Biogeography and Bioresource in Arid Land, Xinjiang Institute of Ecology and Geography, Urumqi 830011, China
3. The Specimen Museum of Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Urumqi 830011, China
4. Institute of Botany, Academy Science Republic of Uzbekistan, Tashkent 100126, Uzbekistan