1. School of Forestry, Northeast Forestry University, Harbin 150040, China
2. Northeast Asia Biodiversity Research Center, Harbin 150040, China
3. School of Geography and Tourism, Harbin University, Harbin 150040, China
4. Folia Multidimensional Innovate Lab, Anshan 114000, China
5. State Key Laboratory of Vegetation and Environmental Change, Institute of Botany, Chinese Academy of Sciences, Beijing 150093, China