1. Department of Plant Science, Faculty of Science, Mahidol University, Bangkok 10400, Thailand
2. Meise Botanic Garden, Nieuwelaan 38, 1860 Meise, Belgium
3. Department of Biology, KU Leuven, 3001 Leuven, Belgium
4. Leuven Plant Institute, KU Leuven, 3001 Leuven, Belgium