1. German Centre for Marine Biodiversity Research DZMB, Senckenberg am Meer, Südstrand 44, 26382 Wilhelmshaven, Germany
2. Department of Biology, Faculty of Mathematics and Natural Sciences, Biozentrum Köln, University of Cologne, Zülpicher Straße 47b, 50674 Cologne, Germany
3. Faculty of Mathematics and Natural Sciences, Institute of Zoology, Poppelsdorfer Schloss, Rheinische Friedrich-Wilhelms-Universität Bonn, Meckenheimer Allee 169, 53115 Bonn, Germany
4. WG Plant Biodiversity and Evolution, School V, Institute of Biology and Environmental Sciences (IBU), Carl von Ossietzky University of Oldenburg, Carl-von-Ossietzky-Straße 9-11, 26111 Oldenburg, Germany