1. College of Biological Sciences and Technology, Beijing Forestry University, Beijing 100083, China
2. Key Laboratory of Bio-Resources and Eco-Environment of Ministry of Education, College of Life Sciences, Sichuan University, Chengdu 610065, China
3. National Engineering Research Center of Tree Breeding and Ecological Restoration, Beijing Forestry University, Beijing 100083, China
4. Sichuan-Chongqing Key Laboratory of Characteristic Biological Resources Research and Utilization, Chengdu 610065, China