1. Cancer Biology Graduate Program, University of Iowa, Iowa City, IA 52242, USA
2. Medical Scientist Training Program, Carver College of Medicine, University of Iowa, Iowa City, IA 52242, USA
3. Holden Comprehensive Cancer Center, University of Iowa, Iowa City, IA 52242, USA
4. Department of Neuroscience and Pharmacology, Carver College of Medicine, University of Iowa, Iowa City, IA 52242, USA
5. Department of Pathology, Carver College of Medicine, University of Iowa, Iowa City, IA 52242, USA