1. Key Laboratory of Environment and Ecology (Ministry of Education), Ocean University of China, Qingdao 266100, China
2. School of Mathematics, Sun Yat-Sen University, Guangzhou 510275, China
3. Key Laboratory of Marine Eco-Environmental Science and Technology, First Institute of Oceanography, Ministry of Natural Resources, Qingdao 266061, China
4. Laboratory for Marine Ecology and Environmental Science, Laoshan Laboratory, Qingdao 266200, China