1. The Kirkhouse Trust, Unit 6 Fenlock Court, Long Hanborough OX29 8LN, UK
2. Section of Crop & Ecosystem Sciences, Department of Plant Sciences, University of California, 1 Shields Avenue, Davis, CA 95616, USA
3. Departamento de Biología Molecular (Área Genética), Universidad de León, 24071 León, Spain
4. Department of Biology, University of Virginia, Charlottesville, VA 22904, USA