1. State Key Laboratory of Aridland Crop Science, College of Agronomy, Gansu Agricultural University, Lanzhou 730070, China
2. School of Biological and Pharmaceutical Engineering, Lanzhou Jiaotong University, Lanzhou 730070, China
3. The James Hutton Institute, Dundee DD2 5DA, UK
4. International Potato Center (CIP), CIP China Center for Asia Pacific (CCCAP), Beijing 102199, China