1. Plant Breeding Institute, School of Life and Environmental Sciences, Faculty of Science, The University of Sydney, 107 Cobbitty Road, Cobbitty, NSW 2570, Australia
2. Buginyanya Zonal Agricultural Research Institute, National Agricultural Research Organisation, Mbale P.O. Box 1356, Uganda
3. School of Science, Hawkesbury Campus, Western Sydney University, Bourke Street, Richmond, NSW 2753, Australia