1. Institute of Plant Ecology, Justus Liebig University Giessen, 35392 Giessen, Germany
2. Laboratory of Extremophile Plants, Center of Biotechnology of Borj Cedria, Hammam-Lif 2084, Tunisia
3. Faculty of Mathematical, Physical and Natural Sciences of Tunis, University of Tunis El-Manar, Tunis 1068, Tunisia