1. Doctoral Program in Biological and Environmental Sciences, SisBio, Herbario UPTC, ViE-DIN, Universidad Pedagógica y Tecnológica de Colombia, Campus Universitario, Tunja 150003, Colombia
2. Department of Botany and Plant Physiology, Faculty of Sciences, Universidad de Málaga, 29010 Málaga, Spain
3. Botany Unit, Department of Pharmacology, Pharmacognosy and Botany, Faculty of Pharmacy, Complutense University, 28040 Madrid, Spain
4. Harvard University Herbaria, Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138-2094, USA
5. Grupo Evolución y Sistemática Tropical, Department of Biology and Chemistry, Universidad de Sucre, Sincelejo 700001, Colombia