1. Department of Biology, Faculty of Science and Arts, King Khalid University, Mohail 61321, Assir, Saudi Arabia
2. Soil Improvement and Conservation Research Department, Soils, Water, and Environment Research Institute (SWERI), Agriculture Research Center (ARC), Giza 12112, Egypt
3. Soil Microbiology Research Department, Soils, Water, and Environment Research Institute (SWERI), Agriculture Research Center (ARC), Giza 12112, Egypt