1. New Valley Vertebrate Palaeontology Centre, New Valley University, Kharga City 172201, Egypt
2. Department of Geology, Faculty of Science, Ain Shams University, Cairo 11566, Egypt
3. Department of Biological Sciences, MacEwan University, Edmonton, AB T5J 4S2, Canada
4. Department of Biology and Biochemistry, University of Bath, Bath BA2 7AY, UK
5. Department of Geology, Faculty of Science, Cairo University, Giza 12613, Egypt