1. School of Natural Sciences, Bangor University, Bangor LL57 2DG, UK
2. Radboud Institute for Biological and Environmental Sciences (RIBES), Radboud University (RU), Heyendaalseweg 135, 6525 AJ Nijmegen, The Netherlands
3. PBL Netherlands Environmental Assessment Agency, Bezuidenhoutseweg 30, 2594 AV The Hague, The Netherlands
4. Environmental Systems Analyses Group, Wageningen University & Research (WUR), Droevendaalsesteeg 3, 6708 PB Wageningen, The Netherlands