1. College of Horticulture, Qingdao Agricultural University, Qingdao 266109, China
2. China Engineering Laboratory of Genetic Improvement of Horticultural Crops of Shandong Province, Qingdao Agricultural University, Qingdao 266109, China
3. College of Life Sciences, Qingdao Agricultural University, Qingdao 266109, China
4. Academy of Dongying Efficient Agricultural Technology and Industry on Saline and Alkaline Land in Collaboration with Qingdao Agricultural University, Dongying 257300, China