1. South China Sea Environment Monitoring Center, State Oceanic Administration, Guangzhou 510300, China
2. Key Laboratory of Marine Environmental Survey Technology and Application, Ministry of Natural Resources, Guangzhou 510300, China
3. Key Laboratory of Marine Ecosystem Dynamics, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou 310012, China
4. College of Light Industry and Food, Zhongkai University of Agriculture and Engineering, Guangzhou 510225, China