1. Third Institute of Oceanography, Ministry of Natural Resources, Xiamen 361005, China
2. School of Life Science, Central China Normal University, Wuhan 430079, China
3. Fisheries College, Jimei University, Xiamen 361021, China
4. South China Sea Fisheries Research Institute, Chinese Academy of Fisheries Sciences, Guangzhou 510300, China
5. Fujian Provincial Key Laboratory of Marine Fishery Resources and Eco-Environment, Xiamen 361021, China