1. Chengde Academy of Agriculture and Forestry Sciences, Chengde 067000, China
2. Chengde Agriculture and Rural Affairs Bureau, Chengde 067000, China
3. Forestry Biotechnology Hunan Key Laboratories, Central South University of Forestry and Technology, Changsha 410004, China
4. Key Laboratory of Biodiversity Conservation of National Forestry and Grassland Administration, Ecology and Nature Conservation Institute, Chinese Academy of Forestry, Beijing 100091, China