The Imitation Game: In Search for Brachycera in the Triassic

Abstract

The richest assemblage of the Triassic Diptera has been described from the famous Konservat-Lagerstätte Grès à Voltzia (Upper Buntsandstein) in the northern Vosges Mountains in France, dated as Early Anisian. A re-examination of the holotypes and additional material from the type locality allows for the establishment of Vogerhyphus gen. nov. and erection of the Vogerhyphinae subfam. nov. for Vymrhyphus blagoderovi Krzemiński and Krzemińska, 2003 and Vogerhyphus krzeminskorum sp. nov. (Protorhyphidae), and a new monotypic family Galliidae fam. nov. for Gallia alsatica Krzemiński and Krzemińska, 2003, originally described as Rhagionidae based on its wing venation. Galliidae fam. nov. is characterized by its closed cua cell and long moniliform antenna with 14-segmented flagellum and is hypothesized to belong to the stem-group Brachycera, along with the Late Triassic Prosechamyiidae. The process of brachycerization in the Diptera evolution is briefly discussed.