1. College of Life Sciences, School of Marine Science and Engineering, Qingdao Agricultural University, Qingdao 266109, China
2. CAS and Shandong Province Key Laboratory of Experimental Marine Biology, Center of Deep-Sea Research, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
3. College of Marine Science, University of Chinese Academy of Sciences, Beijing 100049, China
4. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266071, China
5. Key Laboratory of Breeding Biotechnology and Sustainable Aquaculture, Chinese Academy of Sciences, Qingdao 266071, China