1. CAS Key Laboratory of Marine Ecology and Environmental Sciences, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
2. Laoshan Laboratory, Qingdao 266237, China
3. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao 266071, China
4. Yellow Sea and East Sea Buoy Observation Station, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
5. University of Chinese Academy of Sciences, Beijing 100049, China