1. Discipline of Geography, University of KwaZulu-Natal, Westville Campus, PB X54001, Durban 4000, South Africa
2. School of Life Sciences, University of KwaZulu-Natal, Westville Campus, PB X54001, Durban 4000, South Africa
3. Department of Natural Resources, Sabaragamuwa University, P.O. Box. 2, Belihuloya 70140, Sri Lanka