1. National Institute of Ecology, Seocheon 33657, Republic of Korea
2. Department of Environmental Science and Ecological Engineering, Graduate School, Korea University, Seoul 02841, Republic of Korea
3. Center for Endangered Species, National Institute of Ecology, Yeongyang 36531, Republic of Korea
4. School of Integrative Plant Science, Cornell University, Ithaca, NY 14850, USA
5. Forest Technology and Management Research Center, Pocheon-si 11186, Republic of Korea