1. College of Veterinary Medicine, Yangzhou University, Yangzhou 225009, China
2. Department of Diagnostic Medicine & Pathobiology, Kansas State University, 1800 Denison Avenue, Manhattan, KS 66506, USA
3. Comparative Medicine Research Institute, Yangzhou University, Yangzhou 225009, China
4. Jiangsu Co-Innovation Center for Prevention and Control of Important Animal Infectious Diseases and Zoonoses, Yangzhou 225009, China