1. Key Laboratory of Mariculture, Ministry of Education, College of Fisheries, Ocean University of China, Qingdao 266003, China
2. Department of Marine and Coastal Resources, Lak Si, Bangkok 10210, Thailand
3. Research Center for Coastal Zone Science and Marine Development Strategy, The First Institute of Oceanography, Ministry of Natural Resources, No. 6, Xianxialing Road, Qingdao 266061, China