1. School of Biology, Faculty of Science and Technology, Nakhon Ratchasima Rajabhat University, Nakhon Ratchasima 30000, Thailand
2. National Institute of Biology, 1000 Ljubljana, Slovenia
3. Applied Taxonomic Research Center, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand
4. Laboratory of Biodiversity and Environmental Management, International College, Khon Kaen University, Khon Kaen 40002, Thailand