1. NARO Kyushu Okinawa Agricultural Research Center, Suya 2421, Koshi, Kumamoto 861-1192, Japan
2. Department of Agriculture, Tokai University, Toroku 9-1-1, Higashi-ku, Kumamoto 862-8652, Japan
3. Range Cattle Research and Education Center, University of Florida, Ona, FL 33865, USA